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Additionally, excessive electron flow in the photosynthetic transport chain, particularly between photosystem II (PSII) and PSI, increases generations of reactive oxygen species (ROS), including superoxide (O 2 ), hydrogen peroxide (H 2 O 2 ), and methylglyoxal (MG)

-Ketoglutarate plays an essential role in the production of glutamate and -aminobutyric acid (GABA) and contributes to the scavenging of reactive oxygen species

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Loss of the catalytic activity was considered to be attributed to both the absence of glycosylation and the failure of the tetramer