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IEC6 cells were subjected to treatment with metformin alone, post-metformin treatment with the AMPK inhibitor (compound C), or rotenone, a known complex 1 inhibitor 42,43 , alone

For that, the -glutamate-cysteine ligase (GshA) condenses the carboxyl group at the -position of glutamate with the amino group of cysteine, creating the dipeptide -glutamyl-cysteine